The former couple, who called it quits right before the holidays in 2012, are reuniting for Marvel's Guardians of the Galaxy, an action flick that follows a pilot and his team of aliens as they battle cosmic threats, according to Deadline.
This is the second film Cooper and Saldana have starred in together, with their romance kicking off in 2010 after meeting on the set of their drama flick, The Words.
Both of the actors have moved on since their split, with Cooper romancing model Suki Waterhouse and Saldana linked to Italian artist Marco Perego.
It's not clear if there will be any awkward moments for the duo though, considering Cooper will only be voicing his character Rocket Raccoon in the film, rather than physically acting in it.
There's still the matter of the promo tour though, once the movie nears its premiere date.
Guardians of the Galaxy is set to be released Aug. 1, 2014.