Looks like another Hollywood couple has apparently called it quits.
The former lovebirds were first romantically linked in July 2012 while filming The Mortal Instruments: City of Bones in Toronto and they are currently on a nationwide press tour promoting the flick.
In fact, they were seen holding hands on Friday while leaving a photo call at Franklin Mills Mall in Philadelphia, just days before reportedly pulling the plug on their romance.
Despite the breakup, a source tells Us Collins and Bower remain "very good friends," and the twosome have apparently been able to maintain an amicable relationship as they continue to make the promotional rounds.
It wasn't long ago that the Mirror Mirror star was gushing to Glamour U.K. over her former beau, telling the mag the 24-year-old actor is "edgy, witty and charming."
She also admitted she understood why so many costars fall love on set, seemingly touching on the spark she and Jamie shared at their first meeting.
"People are cast because they have chemistry," the 24-year-old beauty explained. "You're spending 24-hour days together and it's intense…plus you really get to know the best and worst sides of people—the rawest sides of people. You're perfect for the role because you have that connection with each other and sometimes that translates into more."
And while she had no trouble commenting on her costar, she refused to divulge details about their relationship.
"I entered into this business knowing your private life can't always remain private. I don't feel the need to profess anything publicly or confirm something that in a normal situation you wouldn't have to speak out about," she said before adding. "If I'm out and I'm photographed with someone, then that is what it is. We took that chance and… whatever. It is what it is."
Lily's rep had "no comment" when asked about the breakup report.
Prior to Collins, Jamie was engaged to his Harry Potter and the Deathly Hallows: Part 1 costar Bonnie Wright. The private pair quietly called it quits in June 2012, just one month before he and Lily were spotted getting cozy offset.
Since the split, Jamie has been spotted out with Ashley Greene—his costar from the Twilight flicks.
Are you sad to hear Lily and Jamie have split? Tell us in the comments!